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David Seward Richard

Curriculum Vitae

• Work Address: Dept of Biology, Susquehanna University, Selinsgrove, PA 17870-1164
• Telephone: (570) 372-4206, Fax: (570) 372-2745.
• e-mail: richard@susqu.edu.

Education and work experience.

• 2004. Professor of Biology. Susquehanna University, Selinsgrove, PA 17870.
• 2001-Present. Chair: Biology Department, Susquehanna University
• 2000 -Present Associate Professor of Biology. Susquehanna University, Selinsgrove, PA 17870. .
• 1993-2000 Assistant Professor of Biology. Susquehanna University, Selinsgrove, PA 17870. .Tenure approved May, 1999.
• 1991-1993. University of North Carolina, Dept. of Biology, Chapel Hill, NC 27599-3280. Research Associate/Instructor.
• 1987- 1993 Post Doctoral Research Associate. Work area: Control of Juvenile hormone production in Drosophila melanogaster. University of North Carolina, Dept. of Biology, Chapel Hill, NC 27599-3280.
• 1990 University of North Carolina, Program in Molecular Biology and Biotechnology. Nucleic Acid Techniques course. Jan. 29 to Feb. 16.
• 1983-1987 University of Edinburgh, Dept. of Zoology, Edinburgh UK. Ph.D. Insect endocrinology. Thesis title, "Endocrinology of photoperiodic diapause induction in two species of Diptera."
• 1982-1983 University of Dundee, Dept. of Biology, Dundee UK. Research assistant, non-degree. Work area, Cyanobacterial toxins.
• 1981-1982 University of Dundee, Dept. of Biochemical Medicine, Ninewells Hospital, Dundee, UK. M.Sc. Analytical Biochemistry.
• 1977-1981 University of Stirling, Stirling UK. B.Sc. Major in Biology.

Teaching experience.

• June 1999/2001/2003/2005. Focus Australia study tour program director.
• 1999/2001/2003/2005. Biological Issues in Australia. Seminar course
• 1995-2004 Cell and Organismal Biology. lectures and laboratories, . Introductory level.
• 1998/2000 Human Physiology. Lecture and laboratories
• 2000/2004 Exercise and extreme physiology. Lecture and laboratories
• 1993-present Undergraduate research projects. Supervisor .
• 1994 Animal Diversity, lectures and laboratories, Introductory level.
• 1994-7 Endocrinology , lectures and laboratory/supervised research Upper level.
• 1994-2004 Cell Biology, lectures Cell Biology Lab, . Upper level.
• 1993-2004 Animal Physiology lecture Animal Physiology Lab . Upper level.
• 1993 Biology of AIDS Seminar course, (9 students). Upper level.
• 1993 Introductory Biology, Two laboratory sections, . Introductory level.
• 1993 Introductory Biology, Instructor. 45 lectures, . Introductory level. Biology Dept. UNC-CH.
• 1991/92/93 Comparative Animal Physiology. Instructor., 45 lectures,. Upper level. Biology Dept. UNC-CH.
• 1990-93 Undergraduate research projects Co-supervisor.UNC-CH.
• 1988 Diapause endocrinology Guest lecturer in Clocks and Photoperiodism course; Biology Dept. UNC-CH.
• 1986/87 Undergraduate research projects Co-supervisor. Edinburgh University.
• 1985/86 Clocks and Photoperiodism Teaching assistant. Zoology Dept, Edinburgh University.

Service etc.

• Panel member: Human and Animal Biology: Developmental and Reproductive Biology study section NASA Life and Biomedical Sciences and Applications Division-wide Review Project 1994 July 6 to 8, 1994. Reston Va. Coordinated by the American Institute of Biological Sciences (AIBS). This panel reviewed grant proposals aimed at placing life science experiments on both the Space Shuttle and the Russian MIR space station.
• October 27/28, 1994 National Institutes of Health. Tropical Medicine and Parasitology Study section (TMPSS) Panel member:
• March 2/3, 1995, National Institutes of Health (TMPSS).
• November 2/3, 1995, National Institutes of Health (TMPSS).
• Reviews for Archives of Insect Biochemistry and Physiology. and the Journal of Insect Physiology and European Journal of Entomology, Science, Proceedings of the National Academy of Science as requested by editorial board members.
• Member Pennsylvania Academy of Science; Beta Beta Beta Biological Honor Society; AAAS.
• Susquehanna University service: Member of various University Committees. These include Library, Academic Honesty, Computer Head Search Committee (1996, 2000), Faculty Affairs (1998-2000), Strategic Planning (1997-98), University Council (1998-2002), Chair Middle States accreditation taskforce, 2002-03.
• Member of Board of Directors, Susquehanna University. 2000-2006.
• Susquehanna University Radiation Safety Officer 1995-present. SU Radiation Safety manual

Grants and Awards.

• The role of ecdysteroids and juvenile hormones in adult female reproductive development in Drosophila melanogaster. Susquehanna University Research Grant. January 1994. $5005. PI
• An analysis of hormone binding in reproductively active Drosophila melanogaster females. Susquehanna University Research Grant. January 1996. $4986. PI
• Endocrine control of vitellogenesis in Drosophila. National Institutes of Health, Academic Research Enhancement Award. June 1997. $106,104. Abstract. PI.
• 1999-2000 John C. Horn Award recipient for distinguished scholarship and service to Susquehanna University. The Horn Award was created to recognize and honor Dr. John C. Horn, who served for 24 years as a member of the Board of Directors at Susquehanna University. Press Release
• Acquisition of a confocal microscope to support research in cell and molecular biology at Susquehanna University. National Science Foundation. October, 2001. $129,552. (Co-Pi)
• Endocrine control of vitellogenesis in Drosophila. National Institutes of Health. Academic Research Enhancement Award. March 2002. $127,609. Abstract. PI

Publications.

Muntz,W.R.A. and Richard,D.S. (1982) Photomechanical movements in the trout retina following brief flashes of light. Vision Research,22,529-530.

Richard,D.S., Beattie,K.A. and Codd,G.A. (1983) Toxicity of cyanobacterial blooms from Scottish freshwaters. Environmental Technology Letters, 4, 377-382.

Richard,D.S. and Saunders,D.S. (1986) In vitro activation of prothoracic glands from diapause and non-diapause destined Sarcophaga argyrostoma and Calliphora vicina. In, Insect Neurochemistry and Neurophysiology. Eds. Borkovec,A.B. and Gelman,D.B. Humana Press, pp 343-346.

Richard,D.S., Saunders,D.S., Egan,V. and Thomson,R.C.K. (1986) The timing of larval wandering and puparium formation in the flesh-fly Sarcophaga argyrostoma. Physiological Entomology, 11, 53-60.*

Richard,D.S., Warren,J.T., Saunders,D.S. and Gilbert,L.I. (1987) Haemolymph ecdysteroid titres in diapause and non-diapause destined larvae and pupae of Sarcophaga argyrostoma. J. Insect Physiology, 33, 115-122.

Richard,D.S. and Saunders,D.S. (1987) Prothoracic gland function in diapause and non-diapause destined Sarcophaga argyrostoma and Calliphora vicina. J. Insect Physiology, 33, 385-392.

Saunders,D.S., Kenny,N.A.P. and Richard,D.S. (1988) The blowfly, Calliphora vicina: Some problems in photoperiodism and diapause endocrinology. In, Endocrinological Frontiers in Physiological Insect Ecology. Eds. Sehnal,F., Zabza,A. and Denlinger,D.L. Wroclaw Technical University Press, Wroclaw. pp, 293-308.

Richard,D.S., Applebaum,S.W., Sliter,T.J., Baker,F.C., Schooley,D.A., Reuter,C.C. Henrich,V.C. and Gilbert,L.I. (1989) Juvenile hormone biosynthesis in vitro by the ring gland of Drosophila melanogaster: A putative new juvenile hormone in the higher Diptera. Proceedings of the National Academy of Science, USA. 86, 1421-1425

Richard,D.S., Applebaum,S.W. and Gilbert,L.I. (1989) Developmental regulation of juvenile hormone biosynthesis by the ring gland of Drosophila melanogaster. J. Comparative Physiology B. 159,383-387.

Richard,D.S., Applebaum,S.W. and Gilbert,L.I. (1990) Allatostatic regulation of juvenile hormone production in vitro by ring glands of Drosophila melanogaster . Molecular and Cellular Endocrinology. 68, 153-161.

Altaratz,M., Segal,D., Richard,D.S., Gilbert,L.I. and Applebaum,S.W. (1990) Juvenile hormone production by wild type and apterous4 mutant Drosophila melanogaster corpora allata in vitro. In, Insect Neurochemistry and Neurophysiology 1989. Eds. A.B. Borkovec and E.P. Masler. The Humana Press, pp 333-336.

Saunders,D.S., Richard,D.S. Applebaum,S.W., Ma,M. and Gilbert,L.I. (1990)Photoperiodic diapause in Drosophila melanogaster: A block to the juvenile hormone regulation of ovarian maturation. General and Comparative Endocrinology. 79,174-184.

Baker,F.C., Reuter,C.C., Tsai,L.W., Brindle,P.A., Richard,D.S., Tobe,S.S. and Schooley,D.A. (1990) Use of microderviatization techniques in combination with thin-layer chromatography, liquid chromatography and gas chromatography-mass spectrometry for investigation of juvenile hormones and related compounds. In, Chromatography and isolation of insect hormones and pheromones. Eds. McCaffery, A.R. and Wilson, I.D. Plenum Press, New York, pp 19-30.

Altaratz,M., Segal,D., Richard,D.S., Gilbert,L.I. and Applebaum,S.W. (1991) Regulation of juvenile hormone synthesis in wild-type and apterous mutant Drosophila. Molecular and Cellular Endocrinology. 81,205-216.

Richard, D.S. and Gilbert,L.I. (1991) Reversible juvenile hormone inhibition of ecdysteroid and juvenile hormone synthesis by the ring gland of Drosophila melanogaster. Experientia. 47,1063-1066.

Gilbert,L.I. and Richard,D.S. (1991) The juvenile hormone of Drosophila melanogaster. Am. Chem. Soc. Abstr. Pap. 202, 1-2.

Segal,D., Altaratz,M., Richard,D.S., Gilbert,L.I. and Applebaum,S.W. (1991) Developmental regulation of juvenile hormone synthesis in wild-type and apterous mutant Drosophila melanogaster. Phytoparasitica, 19,4. 333-334

Sliter,T.J., Warren,J.L., Richard,D.S. and Gilbert,L.I. (1992) A new Drosophila mutant with novel effects on ecdysteroid-regulated development. Abs. Ecdysone Workshop 10. p71.

Kenny, N.A.P., Richard,D.S., Bradley,H.K. and Saunders,D.S. (1992) Photoperiodic sensitivity and diapause induction during ovarian, embryonic and larval development of the flesh fly Sarcophaga argyrostoma. Journal of Biosciences. 17,241-252.

Richard, D.S. Arnim,A.E. and Gilbert, L.I. * (1993) A reappraisal of the hormonal regulation of larval fat body histolysis in female Drosophila melanogaster. Experientia. 49,150-156. *

Billings,J., Kern,A., Peeler,M. and Richard, D.S. (1995). Teratogenic effects of retinoic acid in sea urchin development. Abs. J. Pennsylvania Academy of Science. 68,168-169. *

Jayawardena,J. McGonnigle,J. and Richard,D.S. (1996) A mutational examination of hormonal regulation of vitellogenesis in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 69,150. * Jayawardena abstract

Purcell,N.H. and Richard,D.S. (1996) Ecdysteroid and juvenile hormone production in diapausing female Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 69,160. * Purcell abstract

Robbins,K.M., Evans,D.M., Richard,D.S. and Peeler,M.T. (1996) Retinoic acid affects vegetal plate formation in sea urchin embryos. Abs. J. Pennsylvania Academy of Science. 69,161-162. * Robbins abstract

Hollinshead D. M. and. Richard.D.S. (1997) Immunofluorescent localization of clathrin to Drosophila melanogaster oocytes. Abs. J. Pennsylvania Academy of Science. 70,174-175. * Hollinshead abstract

Watkins,W., Serafin,R. and Richard, D.S. (1997) Ecdysteroids stimulate vitellogenesis in ovaries from diapausing female Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 70,198. * Watkins abstact

Flango,J.K., Richard, D.S. and Peeler,M.T. (1997) The effects of all-trans-retinoic acid on morphology and gene expression in the sea urchin species Lytechinus pictus and Lytechinus variagatus. Abs. J. Pennsylvania Academy of Science. 70,172. *

Richard,D.S., Watkins,N.L., Serafin,R.B., and Gilbert L.I. (1998) Ecdysteroids regulate yolk protein uptake by Drosophila melanogaster oocytes. Journal of Insect Physiology. 44,637-644* JIP abstract

Richard,D.S., Watkins,N.L., Serafin,R.B., Jones,J.M. and Hollinshead D.M. (1998) Ecdysteroids drive the uptake of yolk proteins by oocytes from Drosophila melanogaster. 1998 Keystone meeting, "Towards the Genetic manipulation of insects." * Keystone abstract

Gilbert,M. and Richard,D.S. (1998) Clathrin and adaptin levels in ooctyes from Drosophila melanogaster measured by immunofluorescence and western blotting. Abs. J. Pennsylvania Academy of Science. 71,174.* Gilbert abstract

Jones,J.M., Barbarito,M.R. and Richard,D.S. (1998) Ecdysteroids drive the uptake of yolk proteins by oocytes from diapausing Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 71,180.* Jones abstract

Stacy Cerula, Adam Horst, J. Paul Detweiler, Jennifer M. Jones and David S. Richard. (1999) Ecdysteroid and Juvenile Hormone Binding Proteins are regulated during Vitellogenic Development in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 72,155. *Cerula

Brandy Crum, Matthew Gilbert and David S. Richard. (1999) Immunofluorescent Localization of Yolk Protein Receptors and Adaptin in Developing Oocytes of Drosophila melanogaster.Abs. J. Pennsylvania Academy of Science. 72.157*Crum

Suzanne Schelble* and David S. Richard. (1999) Autonomous termination of diapause by ovaries from Drosophila melanogaster as determined by immunofluorescent localization of adaptin and the yolk protein receptor. Abs. J. Pennsylvania Academy of Science. 72,179. *Schelble

Richard,D.S., Jones,J.M., Crum,B., Gilbert,M., Barbarito,M. and Schelble,S. (1999) Hormonal regulation of yolk protein uptake by vitellogenic oocytes in diapausing and mutant female Drosophila melanogaster. Presented to JH VII. Conference held in Jerusalem, Israel, August, 1999. JH VII abstract

Brannigan, Dana M. , Fischer, Stephen J. and Richard, David S. (2000) Juvenile hormone synthesis by tissues other than the corpus allatum during vitellogenic development in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 73, 149.* Brannigan

Cline, Deborah E. *, Zuretti, Gina M. and Richard, David S. (2000)Adaptin and the Yolk Protein Receptor appear to localize to Follicle Cells in Early Vitellogenic Oocytes in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 73,152 * Cline

David S. Richard, Jennifer M. Jones, Matthew Gilbert, Michael Barbarito, Dana Brannigan and Stephen Fischer. (2000) Hormonal regulation of yolk protein uptake by vitellogenic oocytes in diapausing and mutant female Drosophila melanogaster. Presented to Complex Clocks. International conference in honour of Professor David Saunders. Held Edinburgh, March 20-24, 2000.* Abstract

Richard,D.S., Gilbert,M., Crum,B. Hollinshead,D.M. Schelble,S. and Scheswohl,D. (2001a) Yolk protein endocytosis by oocytes in Drosophila melanogaster: immunofluorescent localization of clathrin, adaptin and the yolk protein receptor. Journal of Insect Physiology.* 47/7,715-723. Abstract. Full pdf download of proof.

Richard, David S., Jennifer M. Jones, Michael R. Barbarito, Stacy Cerula, J. Paul Detweiler, Stephen J. Fisher, Dana M. Brannigan, Danielle M. Scheswohl. (2001b) Vitellogenesis in diapausing and mutant Drosophila melanogaster; further evidence for the relative roles of ecdysteroids and juvenile hormones. Journal of Insect Physiology. 47/8,905-913..* Abstract. Full pdf download of proof.

Brandon M. Zeigler *, Danielle M. Scheswohl and David S. Richard (2002) Ecdysteroids and juvenile hormones may regulate gene expression during vitellogenesis in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 75,128*. Abstract

Michael Pfeiffer *, Michelle Patrick and David S. Richard (2002) Sex Peptide may regulate vitellogenesis in Drosophila melanogaster. Abs. J. Pennsylvania Academy of Science. 75,118* Abstract

Danielle M. Scheswohl, (David S. Richard) (2002) Regulation of Vitellogenesis by ecdysteroids and juvenile hormones in Drosophila melanogaster. National Conference on Undergraduate Research. Whitewater, Wis. April 24-27. p259. *

Brandi Swietkoski and Angela Ellerman (David S. Richard) (2003) Regulation of the insulin-receptor substrate gene (chico) during reproductive-diapause termination in Drosophila melanogaster. National Conference on Undergraduate Research. University of Utah, Salt Lake City. March 13-15. p275.*

Angela Ellerman and Brandi Swietkoski (David S. Richard) (2003) Regulation of the insulin-receptor substrate gene (chico) during reproductive-diapause termination in Drosophila melanogaster. J. Pennsylvania Academy of Science. *

Kathryn E. Herman (David S. Richard) (2003) Microarray analysis of gene expression in Drosophila melanogaster ovaries following treatment with juvenile hormone or ecdysteroids. National Conference on Undergraduate Research. University of Utah, Salt Lake City. March 13-15.. p262. *

Kathryn E. Herman (David S. Richard) (2003) Microarray analysis of gene expression in Drosophila melanogaster ovaries following treatment with juvenile hormone or ecdysteroids. J. Pennsylvania Academy of Science. *

Richard,D.S., Rybczynski,R., Wilson, T.G., Wang,Y., Wayne,M.L., Zhou,Y., Kroll,T.R., Partridge,L. and Harshman,L.G. (2003) Insulin signaling is necessary for vitellogenesis in Drosophila melanogaster irrespective of the roles of juvenile hormone and ecdysteroids.. Journal of Insect Physiology. Submitted August 2004.

Wade Znosko and Julie Merkle (David S. Richard) (2004) Alternate splicing of the insulin-receptor substrate gene (chico) in ovaries during reproductive development in Drosophila melanogaster. National Conference on Undergraduate Research. University of Indiana IUPUI.*

Eric Burghoffer and Kristi Koch (David S. Richard) (2004) Developmental regulation of the insulin-receptor substrate protein gene (chico) during ovarian development and diapause in Drosophila melanogaster. University of Indiana IUPUI.*

* Denotes papers and abstracts published in collaboration with undergraduate students.